He was enamoured with Daphne Garsington. When she disappeared inside the tomb, he dug unceasingly trying to free her. When her body, contaminated with saline drones, finally emerged from the tomb, he rushed to her and was contaminated himself.
Both of them were returned to their normal selves for a short time when River blew up the dam and flooded the valley with salt water. As the saline drones could not process so much water properly, the couple died trying to hold on to each other. (AUDIO: The Boundless Sea)