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The Cyclors arrived on the Overcast's plane of reality because of the Overcast's collective subconscious which created a psychic tether between them. They fed on the bond they had with the Overcast like parasites. (COMIC: The Organ Grinder) The Cyclors raised the Overcast from savages to being a peaceful race. For twelve hundred years the Overcast prospered due to their help. (COMIC: The Then and the Now)
During the Last Great Time War, the Cyclors annihilated the spiral arm of a galaxy. The Daleks were the first to recruit them, as they offered them a sensation they had never experienced: genocide. (COMIC: The Organ Grinder) The War Doctor was sent to deal with them, (COMIC: Pull to Open) accompanied by the Master. (COMIC: The Organ Grinder)
The Cyclors were warped by Alice Obiefune's activation of the Psilent songbox. The energy of the Songbox also touched a Volatix Cabal Dalek mutant, who deformed them further, turning them into the Malignant. (COMIC: Fast Asleep) The Malignant wiped out over fifty generations of the Overcast. (COMIC: The Then and the Now)