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Ruling the Dalek Empire in the 42nd century, this Dalek Emperor commanded an invasion of the Milky Way from the Seriphia Galaxy.

First sending Daleks to obtain the knowledge of the Library on Kar-Charrat, (AUDIO: The Genocide Machine) the Emperor later launched an invasion into the Milky Way. While it successfully conquered much of the galaxy, its real goal was Project Infinity, as it intended to summon Daleks from another reality. When those Daleks turned out to be appalled by genocide, the Emperor was captured by the Earth Alliance, (AUDIO: "Death to the Daleks!", Project Infinity) but transmitted its consciousness into the human, Susan Mendes, to continue the war. (AUDIO: Dalek War: Chapter One) It erased her consciousness and intended to use her form to con the galaxy into lowering its guard; however, a psychic trap caused the remnants of Mendes to awaken and trigger the destruction of all Dalek technology, removing the Emperor from its human host and resulting in the Great Catastrophe. (AUDIO: Dalek War: Chapter Four)