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En-Gula was a priestess of Ishtar at the temple in Kish, and the niece of Puabi. She entered into the service of Ishtar at age seven when her mother died, and became a priestess at age twelve. A year later, Qataka, posing as Ishtar, took over the temple and En-Gula witnessed the strange goings-on there.

When summoned by Ninani in 2700 BC, she reported what she had seen and was asked to help rid the city of the goddess, to which she agreed. En-Gula later encountered the Seventh Doctor and Ace in the temple, and helped them defeat Ishtar. She eventually married Avram. (PROSE: Timewyrm: Genesys)

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