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Nathaniel was the final person to be turned into a vampire as part of Project: Twilight.

On 4 October 1915, Dr William Abberton injected Nathaniel with the Twilight Virus. As Amelia Doory and Reggie Mead escaped from the Forge's facility, they stopped by Abberton's laboratory to kill him. As Doory drank Abberton's blood, Mead untied Nathaniel from the operating table and let him escape with the other vampires.

Sometime in the next century, Nathaniel was found and killed by Abberton, who had survived his death by becoming a vampire. (AUDIO: Project: Twilight)